Troglophilus spinulosus Chopard, 1921, Di Russo et al. 2014 Phylum: Arthropoda / Class: Insecta / Order: Orthoptera / Family: Rhaphidophoridae
Cave references
Gasparo, F. 2003. Tre campagne biospeleologiche a Creta. Progressione 48:59-62.
Species references
Benda, P., Georgiakakis, P., Dietz, C., Hanak, V., Galanaki, K., Markantonatou, V., Chudarkova, A., Hulva, P. & Horacek, I. 2009. Bats (Mammalia: Chiroptera) of the Eastern Mediterranean and Middle East. Part 7. The bat fauna of Crete, Greece. Acta Societatis Zoologicae Bohemicae, 72 (3-4): 105-190.
Beron, P. 2016. Faune cavernicole de la Grèce. Maison d'édition Est-West., Sofia. pp. 229.
Bosmans, R. & Chatzaki, M. 2005. A catalogue of the spiders of Greece. A critical review of all spider species cited from Greece with their localities. Newsletter Belg. Arachn. Soc., 20 (2): 1-124.
Bosmans, R., Van Keer, J., Russell-Smith, A., Kronestedt, T., Alderweireldt, M., Bosselaers, J. & De Koninck, H. 2013. Spiders of Crete (Araneae). A catalogue of all currently known species from the Greek Island of Crete. Newsletter Belg. Arachn. Soc., 28 (1): 1-147.
Bosselaers, J. & Henderickx, H. 2002. A new Savignia from Cretan caves (Araneae: Linyphiidae). Zootaxa, 109: 1-8.
Deeleman-Reinhold, C. L. & Deeleman, P. R. 1988. Revision des Dysderinae (Araneae, Dysderidae), les especes mediterraneennes occidentales exceptees. Tijdschr. Ent., 131: 141-269.
Di Russo, C., Rampini, M. & Cobolli, M. 2014. The cave crickets of Greece: a contribution to the study of Southern Balkan Rhaphidophoridae diversity (Orthoptera), with the description of a new species of Troglophilus Krauss , 1879. Biodiversity Journal, 5 (3): 397-420.
Gasparo, F. 2003. Tre campagne biospeleologiche a Creta. Progressione, 48 (1): 59-62.
Gasparo, F. 2006. Indagini faunistiche nelle grotte delle isole di Creta e Lesbos (Grecia). Progressione, 52 (1/2): 58-60.
Georgiakakis P, Kafkaletou Diez A, Salvarina I, Benda P, Billington G, Dietz C, Billington J, Cove D, Davison S, Cooke M, Papadatou E. 2023. The Bats of Greece: An Updated Review of Their Distribution, Ecology and Conservation. Animals. 2023; 13(15):2529. https://doi.org/10.3390/ani13152529
Henderickx, H. 2e3. Neobisium (Omatoblothrus) schawalleri sp. nov., a new troglobitic pseudoscorpion from Crete (Arachnida: Pseudoscorpiones: Neobisiidae). Phegea, 28 (2): 5-10.
Huber B.A. 2020. Revision of the spider genus Hoplopholcus Kulczyński (Araneae, Pholcidae). Zootaxa 4726:1–94.
Karakasi D, Paragamian K, Mylonas M, Vardinoyannis K 2024. What`s crawling in the dark? An annotated list of gastropods in Greek caves.. Subterranean Biology 48: 73-116. https://doi.org/10.3897/subtbiol.48.113383
Schawaller, W. 1985. Liste griechischer Neobisiidae mit neuen Höhlenfunden in Epirus, auf Samos und Kreta (Arachnida, Pseudoscorpiones). Stuttgart. Beitr. Naturk. Ser. A. (Biol.), 386: 1-8.
Schmalfuss, H., Paragamian, K. & Sfenthourakis, S. 2004. The terrestrial isopods (Isopoda: Oniscidea) of Crete and the surrounding islands. Stuttgart. Beitr. Naturk. Ser. A. (Biol.), 662: 1-74.
Sendra A., Antić D., Barranco P., Borko Š., Christian E., Delić T., Fadrique F., Faille A., Galli L., Gasparo F., Georgiev D., Giachino P.M., Kováč L., Lukić M., Marcia P., Miculinić K., Nicolosi G., Palero F., Paragamian K., Pérez T., Polak S., Prieto C. 2020. Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1–138. https://doi.org/10.5852/ejt.2020.591
Citation
Paragamian, K. 2026. Fauna of the cave
Spilaio Doxa, Marathos, Malevizi, Kriti. In Paragamian, K., M. Poulinakis, S. Paragamian and I. Nikoloudakis.
Cave fauna of Greece database - Hellenic Institute of Speleological Research.
Cave data updated 25.01.2020,
Accessed 19.04.2026