Spilaio Kamilari

Higher Geography
Region: Kriti,
Sub-Region: Irakleio,
Municipality: Malevizi,
Closest Settlement: Tylisos
Cave Synonyms
"Coubbedes"
Cave type
Natural
Fauna
  • Alopecosa albofasciata (Brullé, 1832), Roewer 1959, Bosmans & Chatzaki 2005, Bosmans et al. 2013
    Phylum: Arthropoda / Class: Arachnida / Order: Araneae / Family: Lycosidae
  • Amaurobius candia Thaler & Knoflach, 2002, Naumova et al. 2016
    Phylum: Arthropoda / Class: Arachnida / Order: Araneae / Family: Amaurobiidae
  • Aporrectodea caliginosa (Savigny, 1826), Omodeo 1956a, Beron 2016, Szederjesi et al. 2017
    Phylum: Annelida / Class: Clitellata / Order: Crassiclitellata / Family: Lumbricidae
  • Aporrectodea rosea (Savigny, 1826), Omodeo 1956a, Beron 2016
    Phylum: Annelida / Class: Clitellata / Order: Crassiclitellata / Family: Lumbricidae
  • Bathytropa granulata Aubert & Dollfus, 1890, Vandel 1958, Schmalfuss 1972b, Schmalfuss 1979, Beron 2016
    Phylum: Arthropoda / Class: Malacostraca / Order: Isopoda / Family: Bathytropidae
  • Bryocamptus pygmaeus (G.O. Sars, 1863), Chappuis 1956, Beron 2016
    Phylum: Arthropoda / Class: Hexanauplia / Order: Harpacticoida / Family: Canthocamptidae
  • Cordioniscus beroni Vandel, 1968, Vandel 1968, Dalens 1970, Schmalfuss 1972b, Schmalfuss & Erhard 1998, Schmalfuss et al. 2004, Beron 2016
    Phylum: Arthropoda / Class: Malacostraca / Order: Isopoda / Family: Styloniscidae
  • Drassodes lutescens (C. L. Koch, 1839), Bosmans et al. 2013
    Phylum: Arthropoda / Class: Arachnida / Order: Araneae / Family: Gnaphosideae
  • Dysdera crocata C.L. Koch, 1838, Roewer 1959, Brignoli 1979a, Deeleman-Reinhold & Deeleman 1988, Bosmans & Chatzaki 2005, Bosmans et al. 2013
    Phylum: Arthropoda / Class: Arachnida / Order: Araneae / Family: Dysderidae
  • Dysdera sp., Beron 2016
    Phylum: Arthropoda / Class: Arachnida / Order: Araneae / Family: Dysderidae
  • Eupolybothrus litoralis (L. Koch, 1867), Zapparoli 1994
    Phylum: Arthropoda / Class: Chilopoda / Order: Lithobiomorpha / Family: Lithobiidae
  • Holocnemus pluchei (Scopoli, 1763), Roewer 1959, Brignoli 1979a, Bosmans & Chatzaki 2005, Bosmans et al. 2013
    Phylum: Arthropoda / Class: Arachnida / Order: Araneae / Family: Pholcidae
  • Hoplopholcus labyrinthi (Kulczynski, 1903), Roewer 1959, Brignoli 1984, Deltshev 1985, Gasparo 2003, Bosmans & Chatzaki 2005, Bosmans et al. 2013, Beron 2016, Huber 2020
    Phylum: Arthropoda / Class: Arachnida / Order: Araneae / Family: Pholcidae
  • Hypsugo savii (Bonaparte, 1837), Benda et al. 2009, Georgiakakis et al. 2023
    Phylum: Chordata / Class: Mammalia / Order: Chiroptera / Family: Vespertilionidae
  • Lacinius horridus (Panzer 1794), Roewer 1959
    Phylum: Arthropoda / Class: Arachnida / Order: Opiliones / Family: Phalangiidae
  • Latridius minutus (Linnaeus, 1767), Coiffait 1955, Beron 2016
    Phylum: Arthropoda / Class: Insecta / Order: Coleoptera / Family: Latridiidae
  • Lithobius crassipes L. Koch, 1862, Matic 1980, Zapparoli 1994, Zapparoli 2002
    Phylum: Arthropoda / Class: Chilopoda / Order: Lithobiomorpha / Family: Lithobiidae
  • Lithobius cretaicus Matic, 1980, Matic 1980, Zapparoli 1994, Stoev & Beron 2000, Zapparoli 2002, Beron 2016
    Phylum: Arthropoda / Class: Chilopoda / Order: Lithobiomorpha / Family: Lithobiidae
  • Maimuna cretica (Kulczynski, 1903), Roewer 1959, Brignoli 1979b, Bosmans & Chatzaki 2005, Bosmans et al. 2013, Beron 2016
    Phylum: Arthropoda / Class: Arachnida / Order: Araneae / Family: Agelenidae
  • Medon beroni Coiffait, 1970, Coiffait 1969a, Coiffait 1976, Bordoni 1980, Beron 2016, Assing 2017
    Phylum: Arthropoda / Class: Insecta / Order: Coleoptera / Family: Staphylinidae
  • Miniopterus schreibersii (Kuhl, 1817), Benda et al. 2009, Georgiakakis et al. 2023
    Phylum: Chordata / Class: Mammalia / Order: Chiroptera / Family: Miniopteridae
  • Monocyphoniscus bulgaricus (Strouhal, 1939), Vandel 1968, Schmalfuss 1972b, Schmalfuss 1979, Beron 2016
    Phylum: Arthropoda / Class: Malacostraca / Order: Isopoda / Family: Trichoniscidae
  • Myotis blythii (Tomes, 1857), Benda et al. 2009, Georgiakakis et al. 2023
    Phylum: Chordata / Class: Mammalia / Order: Chiroptera / Family: Vespertilionidae
  • Myotis capaccinii (Bonaparte, 1837), Benda et al. 2009, Georgiakakis et al. 2023
    Phylum: Chordata / Class: Mammalia / Order: Chiroptera / Family: Vespertilionidae
  • Myotis emarginatus (Geoffroy, 1806), Benda et al. 2009, Georgiakakis et al. 2023
    Phylum: Chordata / Class: Mammalia / Order: Chiroptera / Family: Vespertilionidae
  • Orthometopon phaleronense (Verhoeff, 1901), Schmalfuss 1972b, Schmalfuss 1979, Beron 2016
    Phylum: Arthropoda / Class: Malacostraca / Order: Isopoda / Family: Agnaridae
  • Ovaliptila lindbergi (Chopard 1957), Chopard 1957, Popov 1984, Willemse 1984, Beron 2016
    Phylum: Arthropoda / Class: Insecta / Order: Orthoptera / Family: Gryllidae
  • Pachyiulus varius (Fabricius, 1781), Lang 1964, Felesaki et al. 2010, Beron 2016
    Phylum: Arthropoda / Class: Diplopoda / Order: Julida / Family: Julidae
  • Porcellio laevis Latreille, 1804, Schmalfuss 1972b, Schmalfuss 1979, Beron 2016
    Phylum: Arthropoda / Class: Malacostraca / Order: Isopoda / Family: Porcellionidae
  • Porcellionides myrmecophilus (Stein, 1859)
    Phylum: Arthropoda / Class: Malacostraca / Order: Isopoda / Family: Porcellionidae
  • Pterostichus sp., Coiffait 1955, Beron 2016
    Phylum: Arthropoda / Class: Insecta / Order: Coleoptera / Family: Carabidae
  • Rhinolophus blasii Peters, 1866, Benda et al. 2009, Georgiakakis et al. 2023
    Phylum: Chordata / Class: Mammalia / Order: Chiroptera / Family: Rhinolophidae
  • Rhinolophus ferrumequinum (Schreber, 1774), Benda et al. 2009, Georgiakakis et al. 2023
    Phylum: Chordata / Class: Mammalia / Order: Chiroptera / Family: Rhinolophidae
  • Rhinolophus hipposideros (André, 1797), Benda et al. 2009, Georgiakakis et al. 2023
    Phylum: Chordata / Class: Mammalia / Order: Chiroptera / Family: Rhinolophidae
  • Scolopendra cretica Attems, 1902, Chamberlin 1956
    Phylum: Arthropoda / Class: Chilopoda / Order: Scolopendromorpha / Family: Scolopendridae
  • Scytodes thoracica (Latreille, 1802), Roewer 1959, Brignoli 1979a, Bosmans & Chatzaki 2005, Bosmans et al. 2013, Beron 2016
    Phylum: Arthropoda / Class: Arachnida / Order: Araneae / Family: Scytodidae
  • Tegenaria ariadnae Brignoli, 1984, Roewer 1959, Brignoli 1979a, Bosmans et al. 2013, Beron 2016
    Phylum: Arthropoda / Class: Arachnida / Order: Araneae / Family: Agelenidae
  • Tegenaria pagana C.L. Koch, 1840, Roewer 1959, Brignoli 1979b, Gasparo 2003, Bosmans & Chatzaki 2005, Bosmans et al. 2013, Beron 2016
    Phylum: Arthropoda / Class: Arachnida / Order: Araneae / Family: Agelenidae
  • Tegenaria parietina (Fourcroy, 1785), Roewer 1959, Brignoli 1979b, Bosmans & Chatzaki 2005, Bosmans et al. 2013, Beron 2016
    Phylum: Arthropoda / Class: Arachnida / Order: Araneae / Family: Agelenidae
  • Trichoniscus lindbergi Vandel, 1958, Schmalfuss et al. 2004, Beron 2016, Dimitriou & Sfenthourakis 2022
    Phylum: Arthropoda / Class: Malacostraca / Order: Isopoda / Family: Trichoniscidae
Cave references
  • Lindberg, K. 1955. Notes sur le grottes de l΄ île de Crète. Fragm. balc. Skopje 1(19):165-174.
  • Platakis, E. 1967. The cave "Kamilari" (CRNr 506). Bull. Soc. Speleol. Grèce 9:12–21.
Species references
  • Assing, V. 2017. On the Staphylinidae of the Greek island Ikaría, with supplementary notes on the fauna of Samos (Coleoptera: Staphylinidae). Koleopterologische Rundschau 87: 89-116
  • Benda, P., Georgiakakis, P., Dietz, C., Hanak, V., Galanaki, K., Markantonatou, V., Chudarkova, A., Hulva, P. & Horacek, I. 2009. Bats (Mammalia: Chiroptera) of the Eastern Mediterranean and Middle East. Part 7. The bat fauna of Crete, Greece. Acta Societatis Zoologicae Bohemicae, 72 (3-4): 105-190.
  • Beron, P. 2016. Faune cavernicole de la Grèce. Maison d'édition Est-West., Sofia. pp. 229.
  • Bordoni, A. 1980. Studi sui Paederinae - III - I Medon Steph. paleartici con descrizione di nuove specie mediterranee (Col. Staphylinidae). Boll. Lab. Ent. agr. "F. Silvestri", 37: 73-125.
  • Bosmans, R. & Chatzaki, M. 2005. A catalogue of the spiders of Greece. A critical review of all spider species cited from Greece with their localities. Newsletter Belg. Arachn. Soc., 20 (2): 1-124.
  • Bosmans, R., Van Keer, J., Russell-Smith, A., Kronestedt, T., Alderweireldt, M., Bosselaers, J. & De Koninck, H. 2013. Spiders of Crete (Araneae). A catalogue of all currently known species from the Greek Island of Crete. Newsletter Belg. Arachn. Soc., 28 (1): 1-147.
  • Brignoli, P. M. 1979. Ragni di Grecia XI. Specie nuove o interessanti, cavernicole ed epigee. Rev. suisse Zool., 86 (1): 181-202.
  • Brignoli, P. M. 1979. Considérations zoogéographiques sur les Araignées cavernicoles de Grèce. Biol. gallo-hellen., 8: 223-236.
  • Brignoli, P. M. 1984. Ragni di Grecia XII. Nuovi dati su varie famiglie (Araneae). Rev. suisse Zool., 91 (2): 281-321.
  • Chamberlin, R. V. 1956. On a collection of chilopods from Crete. Ent. news, 67: 51-53.
  • Chappuis, P. A. 1956. Harpacticoides récoltes en Crète par M. K. Lindberg. Folia Balcanica, 1 (3): 15-18.
  • Chopard, L. 1957. Notes sur les Orthoptères cavernicoles de Crète. Bull. Soc. entomol. Fr., 62 (1-2): 26-28.
  • Coiffait, H. 1955. Sur quelques Coléoptères cavernicoles de Grèce. Notes Biospeol., 10 (2): 205-209.
  • Coiffait, H. 1969. Pselaphides balkaniques. Biol. gallo-hellen., 2 (1): 65-67.
  • Coiffait, H. 1976. Clef de determination des Medon de la région paleartique occidentale avec description d' une espèce cavernicole nouvelle. Ann. Speleol., 31: 229-243.
  • Dalens, H. 1970. Un nouveau représentant du genre Cordioniscus (Isopoda, Oniscoidea, Styloniscidae) récolté en Grèce du Nord. Biol. gallo-hellen., 3 (1): 105-108.
  • Deeleman-Reinhold, C. L. & Deeleman, P. R. 1988. Revision des Dysderinae (Araneae, Dysderidae), les especes mediterraneennes occidentales exceptees. Tijdschr. Ent., 131: 141-269.
  • Deltshev, C. 1985. New data conserning cave spiders (Araneae) in Greece with description of a new Leptonetela (Araneae, Leptonetidae). Acta Zool. Bulg., 27: 41-45.
  • Dimitriou CA , Sfenthourakis S. 2022. An all-inclusive approach: A universal protocol for the successful amplification of four genetic loci of all Onscidea. MethodsX (2022), doi: https://doi.org/10.1016/j.mex.2022.101762
  • Felesaki, I., Stoev, P., Simaiakis, S. & Mylonas, M. 2010. A catalogue of the millipedes of Crete (Myriapoda: Diplopoda). Natura Montenegrina, 9 (3): 357-368.
  • Gasparo, F. 2003. Tre campagne biospeleologiche a Creta. Progressione, 48 (1): 59-62.
  • Georgiakakis P, Kafkaletou Diez A, Salvarina I, Benda P, Billington G, Dietz C, Billington J, Cove D, Davison S, Cooke M, Papadatou E. 2023. The Bats of Greece: An Updated Review of Their Distribution, Ecology and Conservation. Animals. 2023; 13(15):2529. https://doi.org/10.3390/ani13152529
  • Huber B.A. 2020. Revision of the spider genus Hoplopholcus Kulczyński (Araneae, Pholcidae). Zootaxa 4726:1–94.
  • Lang, J. 1964. Diplopoden aus Griechenland, Kreta, der Türkei und Iran aus dem Sammlungen von Dr. K. Lindberg. Acta Soc. Zool. Bohemoslov., 28 (3): 237-246.
  • Matic, Z. 1980. Chilopodes recueillis dans l΄ île Créta. Acta Zool. Bulg., 15: 99-102.
  • Naumova, M., Lazarov, S., Petrov, B. & Deltshev, C. 2016. New faunistic data on the cave-dwelling spiders in the Balkan Peninsula (Araneae). Ecologica Montenegrina, 7: 425-438.
  • Omodeo, P. 1956. Oligocheti dell΄ Indochina e del Mediterraneo Orientale. Mem. Mus. Civ. St. Nat. Verona, 5: 321-336.
  • Popov, A. 1984. Le genre Discoptila Pant. (Orthoptera, Gryllidae). Biol. gallo-hellen., 11 (1): 65-78.
  • Roewer, C. F. 1959. Die Araneae, Solifuga und Opiliones der Sammlungen des Herrn Dr. K. Lindberg aus Griechenland, Creta, Anatolien, Iran und Indien. Göteborgs kungl. vet. vitterh. Samh. Handl., 8 (4): 3-47.
  • Schmalfuss, H. & Erhard, F. 1998. Die Landisopoden (Oniscidea) Griechenlands. 19. Beitrag: Gattung Cordioniscus (Styloniscidae). Stuttgart. Beitr. Naturk. Ser. A. (Biol.), 582: 1-20.
  • Schmalfuss, H. 1972. Die Isopoden von Kreta. Biol. gallo-hellen., 4 (1): 33-60.
  • Schmalfuss, H. 1979. Revidierte Check-List der Landisopoden (Oniscoidea) Griechenlands. Stuttgart. Beitr. Naturk. Ser. A. (Biol.), 331: 1-42.
  • Schmalfuss, H., Paragamian, K. & Sfenthourakis, S. 2004. The terrestrial isopods (Isopoda: Oniscidea) of Crete and the surrounding islands. Stuttgart. Beitr. Naturk. Ser. A. (Biol.), 662: 1-74.
  • Stoev, P. & Beron, P. 2000. On the type specimens of Myriapoda (Chilopoda, Diplopoda) in the collection of the National Museum of Natural History in Sofia.. Arthropoda Selecta 9:91–102.
  • Szederjesi, T., Vavoulidou, E., Chalkia, C., Dányi, L. & Csuzdi, C. 2017. An annotated checklist of earthworms of Greece (Clitellata: Megadrili). Zootaxa, 4272 (1): 57-82.
  • Vandel, A. 1958. Isopodes récoltés dans les grottes de la Crète par le Docteur K. Lindberg. Notes biospeol., 12 (2): 81-101.
  • Vandel, A. 1968. Description d΄ un nouveau représentant de genre Cordioniscus (Crustacés, Isopoda, Oniscoidea, Styloniscidae) suivie de considérations sur les voies de migration de certains lignées d΄ Isopodes terrestres. Ann. Speleol., 23 (3): 621-632.
  • Willemse, F. 1984. Catalogue of the Orthoptera of Greece. Fauna Graeciae I. Hellenic Zoological Society. pp. 275.
  • Zapparoli, M. 1994. Note tassonomiche e faunistiche su Chilopodi della Grecia (Chilopoda). Fragm. Entomol., 26 (1): 11-66.
  • Zapparoli, M. 2002. A catalogue of the centipedes from Greece (Chilopoda). Fragm. Entomol., 34 (1): 1-146.
Citation
Paragamian, K. 2026. Fauna of the cave Spilaio Kamilari, Tylisos, Malevizi, Kriti. In Paragamian, K., M. Poulinakis, S. Paragamian and I. Nikoloudakis. Cave fauna of Greece database - Hellenic Institute of Speleological Research. Cave data updated 23.06.2022, Accessed 03.04.2026