Spilaio Arkoudospilia

Higher Geography
Region: Kentriki Makedonia,
Sub-Region: Pella,
Municipality: Almopia,
Closest Settlement: Loutra Loutrakiou
Cave Synonyms
Cave "A", Agiasma cave, Temna pestera
Cave type
Natural
Protection status
  • GR1240001 - Koryfes Orous Vora (Special Area of Conservation): Law 3937/2011 (OGG/Α 60/31.03.2011), Joint Ministerial Decision 50743/2017 (OGG 4432/B/15.12.17). Check site’s Standard Data Form
  • GR1240008 - Oros Voras (Special Protection Area): Law 3937/2011 (OGG/Α 60/31.03.2011), Joint Ministerial Decision 50743/2017 (OGG 4432/B/15.12.17). Check site’s Standard Data Form
Fauna
  • Alpioniscus gueorguievi Andreev, 2013, Andreev 2013, Beron 2016
    Phylum: Arthropoda / Class: Malacostraca / Order: Isopoda / Family: Trichoniscidae
  • Alpioniscus vejdovskyi (Frankenberger, 1939), Vandel 1946, Rémy 1951, Schmalfuss 1979, Beron 2016
    Phylum: Arthropoda / Class: Malacostraca / Order: Isopoda / Family: Trichoniscidae
  • Dolichopoda remyi Chopard, 1934, Chopard 1940, Willemse 1984, Allegrucci et al. 2011, Di Russo et al. 2014
    Phylum: Arthropoda / Class: Insecta / Order: Orthoptera / Family: Rhaphidophoridae
  • Eupolybothrus macedonicus (Verhoeff, 1943), Verhoeff 1943, Stoev 1997, Zapparoli 2002, Stoev et al. 2010, Beron 2016
    Phylum: Arthropoda / Class: Chilopoda / Order: Lithobiomorpha / Family: Lithobiidae
  • Hussonella remyi Jeannel, 1934, Jeannel 1934, Beron 2016, Fresneda et al. 2024
    Phylum: Arthropoda / Class: Insecta / Order: Coleoptera / Family: Leiodidae
  • Lithobius auritus (Verhoeff, 1943), Verhoeff 1943, Zapparoli 2002, Beron 2016
    Phylum: Arthropoda / Class: Chilopoda / Order: Lithobiomorpha / Family: Lithobiidae
  • Lithobius temnensis Verhoeff, 1943, Verhoeff 1943, Rémy 1951, Zapparoli 2002, Beron 2016
    Phylum: Arthropoda / Class: Chilopoda / Order: Lithobiomorpha / Family: Lithobiidae
  • Meta sp., Beron 2016
    Phylum: Arthropoda / Class: Arachnida / Order: Araneae / Family: Tetragnathidae
  • Psyllipsocus ramburii Selys-Longchamps, 1872, Badonnel 1943, Beron 2016
    Phylum: Arthropoda / Class: Insecta / Order: Psocoptera / Family: Psyllipsocidae
  • Rhinolophus blasii Peters, 1866, Georgiakakis et al. 2023
    Phylum: Chordata / Class: Mammalia / Order: Chiroptera / Family: Rhinolophidae
  • Rhinolophus hipposideros (André, 1797), Georgiakakis et al. 2023
    Phylum: Chordata / Class: Mammalia / Order: Chiroptera / Family: Rhinolophidae
Cave references
  • Rémy, P. 1951. Description de grottes de Macédoine grecque. Ann. Speleol. 6(2-3):107-118.
Species references
  • Allegrucci, G., Trucchi, E. & Sbordoni, V. 2011. Tempo and mode of species diversification in Dolichopoda cave crickets (Orthoptera, Rhaphidophoridae). Molecular Phylogenetics and Evolution, 60 (1): 108-121.
  • Andreev, S. 2013. A new cave species, Alpioniscus gueorguievi n. sp. (Isopoda, Oniscidea, Trichoniscidae) from continental Greece. Acta Zool. Bulg., 65 (3): 297-298.
  • Badonnel, A. 1943. Psocoptères cavernicoles de Macédoine et Herzégovine (Voyages de P. Remy R. Husson et A. Schweitzer).. Bull. Soc. Ent. France, 48: 8-9
  • Beron, P. 2016. Faune cavernicole de la Grèce. Maison d'édition Est-West., Sofia. pp. 229.
  • Chopard, L. 1940. Orthoptères cavernicoles recueillis par P. Remy en Yougoslavie et en Macédoine. Comparaison entre la distribution des Orthoptères et des Coléoptères cavernicoles d΄ Europe. 6eme Congres Intern. Ent. Madrid., Madrid. pp. 509-514.
  • Di Russo, C., Rampini, M. & Cobolli, M. 2014. The cave crickets of Greece: a contribution to the study of Southern Balkan Rhaphidophoridae diversity (Orthoptera), with the description of a new species of Troglophilus Krauss , 1879. Biodiversity Journal, 5 (3): 397-420.
  • Fresneda J, Giachino P M, Salgado J M, Faille A, Bourdeau C, Cieslak A e Ribera I. 2024. A phylogenetic classification of Leptodirini (Coleoptera, Leiodidae, Cholevinae). Memorie della Società Entomologica Italiana, 101(1): 3–936. doi: 10.4081/memoriesei.2024.3.
  • Georgiakakis P, Kafkaletou Diez A, Salvarina I, Benda P, Billington G, Dietz C, Billington J, Cove D, Davison S, Cooke M, Papadatou E. 2023. The Bats of Greece: An Updated Review of Their Distribution, Ecology and Conservation. Animals. 2023; 13(15):2529. https://doi.org/10.3390/ani13152529
  • Jeannel, R. 1934. Bathysciinae recueillis par MM. P. Remy et R. Husson dans le Sandjak de Novi-Pazar et la Macédoine Grecque. Rev. fr. Ent., 1 (2): 89-103.
  • Rémy, P. 1951. Description de grottes de Macédoine grecque. Ann. Speleol., 6 (2-3): 107-118.
  • Schmalfuss, H. 1979. Revidierte Check-List der Landisopoden (Oniscoidea) Griechenlands. Stuttgart. Beitr. Naturk. Ser. A. (Biol.), 331: 1-42.
  • Stoev, P. 1997. A check-list of the centipedes of the Balkan peninsula with some taxonomic notes and a complete bibliography (Chilopoda).. Entomologica scandinavica, Copenhagen, Supplement, 51: 87-105.
  • Stoev, P., Akkari, N., Zapparoli, M., Porco, D., Enghoff, H., Edgecombe, G. D., Georgiev, T. & Penev, L. 2010. The centipede genus Eupolybothrus Verhoeff, 1907 (Chilopoda: Lithobiomorpha: Lithobiidae) in North Africa, a cybertaxonomic revision, with a key to all species in the genus and the first use of DNA barcoding for the group. ZooKeys, 50 (SPEC. ISSUE): 29-77.
  • Vandel, A. 1946. Isopodes terrestres récoltés par M. le Professeur Remy au cours de ses voyages dans les régions balkaniques. Ann. Sci. nat. Zool., 11 (8): 151-194.
  • Verhoeff, K. W. 1943. Über Chilopoden aus westbalkanischen Höhlen.. Zeitschrift für Karst- und Höhlenkunde , 43: 133-152.
  • Willemse, F. 1984. Catalogue of the Orthoptera of Greece. Fauna Graeciae I. Hellenic Zoological Society. pp. 275.
  • Zapparoli, M. 2002. A catalogue of the centipedes from Greece (Chilopoda). Fragm. Entomol., 34 (1): 1-146.
Citation
Paragamian, K. 2026. Fauna of the cave Spilaio Arkoudospilia, Loutra Loutrakiou, Almopia, Kentriki Makedonia. In Paragamian, K., M. Poulinakis, S. Paragamian and I. Nikoloudakis. Cave fauna of Greece database - Hellenic Institute of Speleological Research. Cave data updated 30.04.2020, Accessed 09.03.2026