Pyrenees (Andorra, Spain and France), the Alps from France to Slovenia, Dinaric Alps, Greece including Crete, Corsica and Anatolia through Caucasus to south Iran and Syria
Spilaio Tzani Spilios, Benda et al. 2009, Benda et al. 2019, Georgiakakis et al. 2023 Region: Kriti / Regional unit: Chania / Municipality: Platanias / Closest settlement: Lakkoi
Spilaio Mikro Lavyrinthaki, Hanak et al. 2001, Juste et al. 2004, Benda et al. 2009, Georgiakakis et al. 2023 Region: Kriti / Regional unit: Irakleio / Municipality: Faistos / Closest settlement: Moroni
Spilaio Idaio Antro, Benda et al. 2009, Benda et al. 2019, Georgiakakis et al. 2023 Region: Kriti / Regional unit: Rethymno / Municipality: Anogeia / Closest settlement: Anogeia
Varathro Tafkos sta Petradolakia, Benda et al. 2009, Georgiakakis et al. 2023 Region: Kriti / Regional unit: Rethymno / Municipality: Anogeia / Closest settlement: Anogeia
IUCN Red List
LC - Least Concern
Greece's Red Data Book
EN - Endangered
Protection status
Bern Convention: Appendix II in Council of Europe 1979 - Convention on the conservation of European wildlife and natural habitats / Law 1335/1983, OGG 32/Α/14-03-1983 (Integration of the Bern Convention)
Bonn Convention: Appendix II in Convention on the Conservation of Migratory Species of Wild Animals, 1979 / Law 2719/1999, OGG 106/26-5-1999 (Integration of the Bonn Convention)
Habitats Directive (92/43/EEC): Appendix IV / Joined Ministerial Decision 33318/3028/1998, OGG 1289/Β/1998 (Integration of the Habitats Directive 92/43/EEC)
Law 86/1969, OGG7/Α/1969 (Forest Code modified by Law 2637/1998, OGG 200/A/1998)
Benda, P., Georgiakakis, P., Dietz, C., Hanak, V., Galanaki, K., Markantonatou, V., Chudarkova, A., Hulva, P. & Horacek, I. 2009. Bats (Mammalia: Chiroptera) of the Eastern Mediterranean and Middle East. Part 7. The bat fauna of Crete, Greece. Acta Societatis Zoologicae Bohemicae, 72 (3-4): 105-190.
Benda, P., Ševčík, M., Bego, F., Sachanowicz, K., Spitzenberger, F., Tájek, P., Tájková, P., Uhrin, M. 2019. Bats (Mammalia: Chiroptera) of the Eastern Mediterranean and Middle East. Part 15. The fauna of bats and bat ectoparasites of Albania with a catalogue of bats from the western Balkans in the collection of the National Museum, Prague. Acta Soc. Zool. Bohem. 83: 1–238
Georgiakakis P, Kafkaletou Diez A, Salvarina I, Benda P, Billington G, Dietz C, Billington J, Cove D, Davison S, Cooke M, Papadatou E. 2023. The Bats of Greece: An Updated Review of Their Distribution, Ecology and Conservation. Animals. 2023; 13(15):2529. https://doi.org/10.3390/ani13152529
Hanak, V., Benda, P., Ruedi, M., Homcek, I. & Sofianidou, T. 2001. Bats (Mammalia: Chiroptera) of the Eastern Mediterranean. Part 2. New records and review of distribution of bats in Greece. Acta Societatis Zoologicae Bohemicae, 65: 279-346.
Juste, J., Ibáñez, C., Muñoz, J., Trujillo, D., Benda, P., Karataş, A. & Ruedi, M. 2004. Mitochondrial phylogeography of the long-eared bats (Plecotus) in the Mediterranean Palaearctic and Atlantic Islands. Molecular Phylogenetics and Evolution, 31 (3): 1114-1126.
Citation
Paragamian, K. 2026. Plecotus macrobullaris
Kuzjakin, 1965.
In Paragamian, K., M. Poulinakis, S. Paragamian and I. Nikoloudakis.
Cave fauna of Greece database - Hellenic Institute of Speleological Research.
Species data updated 17.11.2025,
Accessed 17.04.2026