Spilaio Za

Higher Geography

Region: Notio Aigaio,
Sub-Region: Naxos,
Municipality: Naxos & Mikres Kyklades,
Closest Settlement: Filoti

Cave Synonyms

Spilia Aria

Cave type

Natural

Protection status

• GR4220014 - Kentriki and Notia Naxos: Zas and Vigla Eos Mavrovouni and Thalassia Zoni (ormos Karades - Ormos Moutsounas) (Special Area of Conservation): Law 3937/2011 (OGG/Α 60/31.03.2011), Joint Ministerial Decision 50743/2017 (OGG 4432/B/15.12.17). Check site’s Standard Data Form
• GR4220026 - Naxos: Ori Anathematistra, Koronos, Mavrovouni, Zas, Viglatouri (Special Protection Area): Law 3937/2011 (OGG/Α 60/31.03.2011), Joint Ministerial Decision 50743/2017 (OGG 4432/B/15.12.17). Check site’s Standard Data Form

Fauna

• Acanthopetalum hoplites Strasser, 1973, Mauriès & Karamaouna 1984, Stoev et al. 2008, Beron 2016
  Phylum: Arthropoda / Class: Diplopoda / Order: Callipodida / Family: Schizopetalidae
• Aporrectodea caliginosa (Savigny, 1826), Szederjesi 2015, Szederjesi et al. 2017
  Phylum: Annelida / Class: Clitellata / Order: Crassiclitellata / Family: Lumbricidae
• Aporrectodea rosea (Savigny, 1826), Szederjesi 2015, Szederjesi et al. 2017
  Phylum: Annelida / Class: Clitellata / Order: Crassiclitellata / Family: Lumbricidae
• Dendrobaena byblica (Rosa, 1893), Szederjesi 2015, Szederjesi et al. 2017
  Phylum: Annelida / Class: Clitellata / Order: Crassiclitellata / Family: Lumbricidae
• Dolichopoda naxia Boudou-Saltet, 1972, Karny 1930, Boudou-Saltet 1972a, Willemse 1984, Boudou-Saltet 1985, Allegrucci et al. 2011, Rampini et al. 2012, Di Russo et al. 2014, Beron 2016, Allegrucci et al. 2021
  Phylum: Arthropoda / Class: Insecta / Order: Orthoptera / Family: Rhaphidophoridae
• Eiseniella tetraedra (Savigny, 1826), Szederjesi 2015, Szederjesi et al. 2017
  Phylum: Annelida / Class: Clitellata / Order: Crassiclitellata / Family: Lumbricidae
• Hyleoglomeris beroni Mauriès, 1984, Mauriès & Karamaouna 1984, Mauriès 1984b, Stoev & Beron 2000, Thaler 2000, Golovatch 2013a, Makarov et al. 2013, Beron 2016
  Phylum: Arthropoda / Class: Diplopoda / Order: Glomerida / Family: Glomeridae
• Hypsugo savii (Bonaparte, 1837), Georgiakakis et al. 2023
  Phylum: Chordata / Class: Mammalia / Order: Chiroptera / Family: Vespertilionidae
• Lithobius erythrocephalus C.L. Koch, 1847, Matic & Stavropoulos 1988, Zapparoli 2002
  Phylum: Arthropoda / Class: Chilopoda / Order: Lithobiomorpha / Family: Lithobiidae
• Microscolex dubius (Fletscher, 1887), Szederjesi 2015, Szederjesi et al. 2017
  Phylum: Annelida / Class: Clitellata / Order: Crassiclitellata / Family: Acanthodrilidae
• Microscolex phosphoreus (Dugès, 1837), Szederjesi 2015, Szederjesi et al. 2017
  Phylum: Annelida / Class: Clitellata / Order: Crassiclitellata / Family: Acanthodrilidae
• Philedaphia hauseri (Lienhard, 1988), Lienhard 1988a, Lienhard 1998, Lienhard & Smithers 2002
  Phylum: Arthropoda / Class: Insecta / Order: Psocoptera / Family: Protroctopsocidae
• Pipistrellus kuhlii (Kuhl, 1819), Georgiakakis et al. 2023
  Phylum: Chordata / Class: Mammalia / Order: Chiroptera / Family: Vespertilionidae
• Pipistrellus pipistrellus (Schreber, 1774), Georgiakakis et al. 2023
  Phylum: Chordata / Class: Mammalia / Order: Chiroptera / Family: Vespertilionidae
• Plecotus kolombatovici Dulic, 1980, Georgiakakis et al. 2023
  Phylum: Chordata / Class: Mammalia / Order: Chiroptera / Family: Vespertilionidae
• Rhinolophus blasii Peters, 1866, Georgiakakis et al. 2023
  Phylum: Chordata / Class: Mammalia / Order: Chiroptera / Family: Rhinolophidae
• Rhinolophus ferrumequinum (Schreber, 1774), Georgiakakis et al. 2023
  Phylum: Chordata / Class: Mammalia / Order: Chiroptera / Family: Rhinolophidae
• Rhinolophus hipposideros (André, 1797), Georgiakakis et al. 2023
  Phylum: Chordata / Class: Mammalia / Order: Chiroptera / Family: Rhinolophidae
• Schistophallus samius (E. von Martens, 1889), Beron 2016, Karakasi et al. 2024
  Phylum: Mollusca / Class: Gastropoda / Order: Stylommatophora / Family: Oxychilidae
• Tegenaria montiszasensis Bolzern, Burckhardt & Hänggi, 2013, Bolzern 2010, Bolzern et al. 2013
  Phylum: Arthropoda / Class: Arachnida / Order: Araneae / Family: Agelenidae
• Trichoniscus cavernicola Vandel, 1958, Andreev 1986, Beron 2016
  Phylum: Arthropoda / Class: Malacostraca / Order: Isopoda / Family: Trichoniscidae
• Tychobythinus naxius Besuchet, 1993, Besuchet 1993, Hlaváč et al. 2008, Beron 2016
  Phylum: Arthropoda / Class: Insecta / Order: Coleoptera / Family: Staphylinidae

Cave references

• Petrocheilou, A. 1984. The Greek Caves. Ekdotiki Athinon 160pp.
• Petrocheilou, A. 1962. The cave of Zeus “Za ” or Vakchidon (CRNr 705). Bull. Soc. Speleol. Grèce 6:37–42.

Species references

• Allegrucci, G., Trucchi, E. & Sbordoni, V. 2011. Tempo and mode of species diversification in Dolichopoda cave crickets (Orthoptera, Rhaphidophoridae). Molecular Phylogenetics and Evolution, 60 (1): 108-121.
• Allegrucci G, Rampini M, Chimenti C, Alexiou S & Di Russo C. 2021. Dolichopoda cave crickets from Peloponnese (Orthoptera, Rhaphidophoridae): molecular and morphological investigations reveal four new species for Greece. The European Zoological Journal, 88(1): 505-524, DOI: 10.1080/24750263.2021.1902005
• Andreev, S. 1986. Contribution a l΄ étude des isopodes terrestres de la Grèce 3. Sur trois nouvelles donnes sur les Isopodes des Genres Cordioniscus, Alpioniscus et Trichoniscus et nouvelles donnes sur les Isopodes terr. (Oniscoidea). Biol. gallo-hellen., 11 (2): 153-164.
• Beron, P. 2016. Faune cavernicole de la Grèce. Maison d'édition Est-West., Sofia. pp. 229.
• Besuchet, C. 1993. Pselaphides cavernicoles de Grèce (Coleoptera). Biol. gallo-hellen., 20 (1): 223-229.
• Bolzern, A. 2010. Taxonomy and phylogeny of the Tegenaria- Malthonica -complex (Araneae, Agelenidae), using morphological and molecular data.. PHD Thesis. Faculty of Science, University of Basel, Switzerland, 292pp
• Bolzern, A., Burckhardt, D. & Hänggi, A. 2013. Phylogeny and taxonomy of European funnel-web spiders of the Tegenaria-Malthonica complex (Araneae: Agelenidae) based upon morphological and molecular data. Zool. J. Linn. Soc., 168 (4): 723-848.
• Boudou-Saltet, P. 1972. Les Dolichopodes (Orth. Rhaph.) de Grèce VII. Nouvelles espèces du Péloponnèse. Bull. Soc. Hist. Nat. Toulouse, 108 (3-4): 420-425.
• Boudou-Saltet, P. 1985. Etat actuel de nos connaissances sur les orthoptères cavernicoles de Grèce. Biol. gallo-hellen., 10: 259-263.
• Di Russo, C., Rampini, M. & Cobolli, M. 2014. The cave crickets of Greece: a contribution to the study of Southern Balkan Rhaphidophoridae diversity (Orthoptera), with the description of a new species of Troglophilus Krauss , 1879. Biodiversity Journal, 5 (3): 397-420.
• Georgiakakis P, Kafkaletou Diez A, Salvarina I, Benda P, Billington G, Dietz C, Billington J, Cove D, Davison S, Cooke M, Papadatou E. 2023. The Bats of Greece: An Updated Review of Their Distribution, Ecology and Conservation. Animals. 2023; 13(15):2529. https://doi.org/10.3390/ani13152529
• Golovatch, S. I. 2013. Three new species of the millipede genus Hyleoglomeris Verhoeff, 1910 from the Aegean region of Greece (Diplopoda, Glomerida, Glomeridae). Biodiversity Data Journal, 1 (e1000): 1-13.
• Hlaváč, P., Ozimec, R. & Pavićević, D. 2008. Catalogue of the troglobitic Pselaphinae (Coleoptera, Staphylinidae) of the Balkan peninsula, with a key to genera. In Pavićević, D.;Perreau, M. Advances in the studies of the subterranean and epigean fauna of the Balkan Peninsula. Volume dedicated to the memory of Guido Nonveiller. Publisher, Belgrade. pp. 307-328.
• Karakasi D, Paragamian K, Mylonas M, Vardinoyannis K 2024. What`s crawling in the dark? An annotated list of gastropods in Greek caves.. Subterranean Biology 48: 73-116. https://doi.org/10.3897/subtbiol.48.113383
• Karny, H. 1930. Revision der Gryllacriden des Naturhistorischen Museum in Wien einschließlich der Coollektion Brunner von Wattenwyl. Ann. Naturhist. Mus. Wien, 44: 45-198.
• Lienhard, C. and Smithers C.N. 2002. Psocoptera (Insecta) World Catalogue and Bibliography.. Instrumenta Biodiversitatis V. Museum d’histoire naturelle, Genève, Switzerland, pp.xli + 745 pp.
• Lienhard, C. 1988. Three new extra-neotropical species of Troctopsocidae (Insecta : Psocoptera). Journal of Natural History, 22: 575-587.
• Lienhard, C. 1998. Psocoptères Euro-Méditerranéens.. Faune de France 83. Fédération Française des Sociétés de Sciences, Naturelles , xx + 517 pp.
• Makarov SE, Ćurčić BPM, Antić DŽ, Tomić VT, Ćurčić ĆSB, Ilić B, Lučić LR. 2013. A new cave species of the genus Hyleoglomeris Verhoeff, 1910, from the Balkan Peninsula (Diplopoda: Glomerida: Glomeridae). Archive of Biological Sciences, 65(1):341–344
• Matic, Z. & Stavropoulos, G. 1988. Contribution a la connaissance des Chilopodes de Grèce. Biol. gallo-hellen., 14 (1): 33-46.
• Mauriès, J. P. & Karamaouna, M. 1984. Myriapodes-Diplopodes nouveaux et peu connus des îles Naxos, Paros et Antiparos (Cyclades, Grèce). Biol. gallo-hellen., 11 (1): 51-59.
• Mauriès, J. P. 1984. Les Diplopodes récoltés en 1982 par P. Beron et S. Andreev dans l΄ île de Thassos (Grèce). Thassoblaniulus, nouveau genre de Blaniulidae, Blaniulinae. Acta Zool. Bulg., 28: 52-59.
• Rampini, M., Di Russo, C., Taylan, M. S., Gelosa, A. & Cobolli, M. 2012. Four new species of Dolichopoda Bolivar, 1880 from Southern Sporades and Western Turkey (Orthoptera, Rhaphidophoridae, Dolichopodainae). ZooKeys, 201: 43-58.
• Stoev, P. & Beron, P. 2000. On the type specimens of Myriapoda (Chilopoda, Diplopoda) in the collection of the National Museum of Natural History in Sofia.. Arthropoda Selecta 9:91–102.
• Stoev, P., Sierwald, P. & Billey, A. 2008. An annotated world catalogue of the millipede order Callipodida (Arthropoda: Diplopoda).. Zootaxa, 1706:1–50.
• Szederjesi, T. 2015. New earthworm records from various parts of Greece (Oligochaeta: Lumbricidae, Acanthodrilidae, Megascolecidae, Ocnerodrilidae). Opusc. Zool. Budapest, 46 (2): 143-152.
• Szederjesi, T., Vavoulidou, E., Chalkia, C., Dányi, L. & Csuzdi, C. 2017. An annotated checklist of earthworms of Greece (Clitellata: Megadrili). Zootaxa, 4272 (1): 57-82.
• Thaler K. 2000. Über Kugeltausendfüßer aus Griechenland und Zypern (Diplopoda, Glomerida). Entomol. Nachr. Ber. 43:195–201.
• Willemse, F. 1984. Catalogue of the Orthoptera of Greece. Fauna Graeciae I. Hellenic Zoological Society. pp. 275.
• Zapparoli, M. 2002. A catalogue of the centipedes from Greece (Chilopoda). Fragm. Entomol., 34 (1): 1-146.

Citation

Paragamian, K. 2026. Fauna of the cave Spilaio Za, Filoti, Naxos & Mikres Kyklades, Notio Aigaio. In Paragamian, K., M. Poulinakis, S. Paragamian and I. Nikoloudakis. Cave fauna of Greece database - Hellenic Institute of Speleological Research. Cave data updated 25.04.2019, Accessed 09.03.2026