Kokkalis Cave is approximately 3km SE of Vrondero (at 1120 m a.s.l.) and comprises a very large chamber.
Protection status
GR1340001 - Ethnikos Drymos Prespon (Special Protection Area - Special Area of Conservation): Law 3937/2011 (OGG/Α 60/31.03.2011), Joint Ministerial Decision 50743/2017 (OGG 4432/B/15.12.17). Check site’s Standard Data Form
Fauna
Hypsugo savii (Bonaparte, 1837), Papadatou et al. 2011, Papadatou et al. 2012, Georgiakakis et al. 2023 Phylum: Chordata / Class: Mammalia / Order: Chiroptera / Family: Vespertilionidae
Miniopterus schreibersii (Kuhl, 1817), Papadatou et al. 2011, Papadatou et al. 2012, Georgiakakis et al. 2023 Phylum: Chordata / Class: Mammalia / Order: Chiroptera / Family: Miniopteridae
Myotis blythii (Tomes, 1857), Papadatou et al. 2011, Papadatou et al. 2012, Georgiakakis et al. 2023 Phylum: Chordata / Class: Mammalia / Order: Chiroptera / Family: Vespertilionidae
Myotis capaccinii (Bonaparte, 1837), Papadatou et al. 2011, Papadatou et al. 2012, Georgiakakis et al. 2023 Phylum: Chordata / Class: Mammalia / Order: Chiroptera / Family: Vespertilionidae
Myotis myotis (Borkhausen, 1797), Papadatou et al. 2011, Papadatou et al. 2012, Uhrin et al. 2020, Georgiakakis et al. 2023 Phylum: Chordata / Class: Mammalia / Order: Chiroptera / Family: Vespertilionidae
Pipistrellus pipistrellus (Schreber, 1774), Papadatou et al. 2011, Papadatou et al. 2012, Georgiakakis et al. 2023 Phylum: Chordata / Class: Mammalia / Order: Chiroptera / Family: Vespertilionidae
Plecotus austriacus (Fischer, 1829), Papadatou et al. 2011, Papadatou et al. 2012, Uhrin et al. 2020, Georgiakakis et al. 2023 Phylum: Chordata / Class: Mammalia / Order: Chiroptera / Family: Vespertilionidae
Rhinolophus blasii Peters, 1866, Papadatou et al. 2011, Papadatou et al. 2012, Georgiakakis et al. 2023 Phylum: Chordata / Class: Mammalia / Order: Chiroptera / Family: Rhinolophidae
Rhinolophus euryale Blasius, 1853, Papadatou et al. 2011, Papadatou et al. 2012, Georgiakakis et al. 2023 Phylum: Chordata / Class: Mammalia / Order: Chiroptera / Family: Rhinolophidae
Rhinolophus ferrumequinum (Schreber, 1774), Papadatou et al. 2011, Papadatou et al. 2012, Uhrin et al. 2020, Georgiakakis et al. 2023 Phylum: Chordata / Class: Mammalia / Order: Chiroptera / Family: Rhinolophidae
Rhinolophus hipposideros (André, 1797), Papadatou et al. 2011, Papadatou et al. 2012, Uhrin et al. 2020, Georgiakakis et al. 2023 Phylum: Chordata / Class: Mammalia / Order: Chiroptera / Family: Rhinolophidae
Species references
Fehér Z., Szekeres M. 2016. Taxonomic revision of the rock-dwelling door snail genus Montenegrina Boettger, 1877 (Mollusca, Gastropoda, Clausiliidae). ZooKeys 559:1–137.
Georgiakakis P, Kafkaletou Diez A, Salvarina I, Benda P, Billington G, Dietz C, Billington J, Cove D, Davison S, Cooke M, Papadatou E. 2023. The Bats of Greece: An Updated Review of Their Distribution, Ecology and Conservation. Animals. 2023; 13(15):2529. https://doi.org/10.3390/ani13152529
Papadatou, E., Grémillet, X. & Bego, F. 2011. Status survey and conservation action plan for the bats of Prespa. Society for the Protection of Prespa (SPP), Agios Germanos. pp. 170.
Papadatou E., Grémillet X., Bego F. 2012. Status Survey and Conservation Action Plan for Caves and Bats of Prespa. Final report in the framework of the project “Landscape Scale Conservation in the Prespa Lake Basin – Transboundary Species and Habitat Conservation Action Plans.”, pp 54
Uhrin M, Balogova M, Uhrinova R, Vargova V. 2020. Bats of the Prespa Basin in northern Greece: acoustic survey, new records and relationships to light pollution (Chiroptera).. Lynx, n. s. (Praha) 51:147–191
Citation
Paragamian, K. 2025. Fauna of the cave
Spilaio Kokkali, Vrontero, Prespes, Dytiki Makedonia. In Paragamian, K., M. Poulinakis, S. Paragamian and I. Nikoloudakis.
Cave fauna of Greece database - Hellenic Institute of Speleological Research.
Cave data updated 25.05.2018,
Accessed 18.10.2025