Spilaio Agias Triadas

Higher Geography

Region: Sterea Ellada,
Sub-Region: Evvoia,
Municipality: Karystos,
Closest Settlement: Karystos

Cave type

Natural

Protection status

• GR2420012 - Oros Ochi, Paraktia Zoni and Nisides (Special Protection Area): Law 3937/2011 (OGG/Α 60/31.03.2011), Joint Ministerial Decision 50743/2017 (OGG 4432/B/15.12.17). Check site’s Standard Data Form

Fauna

• Acanthopetalum sp., Beron 2016
  Phylum: Arthropoda / Class: Diplopoda / Order: Callipodida / Family: Schizopetalidae
• Amphipyra effusa Boisduval, 1828, Beron 2016
  Phylum: Arthropoda / Class: Insecta / Order: Lepidoptera / Family: Noctuidae
• Ausobskya sp., Beron 2016
  Phylum: Arthropoda / Class: Arachnida / Order: Opiliones / Family: Phalangodidae
• Chthonius sp., Beron 2016
  Phylum: Arthropoda / Class: Arachnida / Order: Pseudoscorpionida / Family: Chthoniidae
• Dolichopoda cassagnaui Boudou-Saltet, 1971, Boudou-Saltet 1971b, Boudou-Saltet 1980, Willemse 1984, Boudou-Saltet 1985, Allegrucci et al. 2011, Rampini et al. 2012, Di Russo et al. 2014, Beron 2016, Allegrucci et al. 2021
  Phylum: Arthropoda / Class: Insecta / Order: Orthoptera / Family: Rhaphidophoridae
• Euscorpius birulai Fet, Soleglad, Parmakelis, Kotsakiozi & Stathi, 2014, Fet et al. 2014, Kovařik & Beron 2015, Beron 2016, Fet et al. 2018, Blasco-Aróstegui & Prendini 2024
  Phylum: Arthropoda / Class: Arachnida / Order: Scorpiones / Family: Euscorpiidae
• Graeconiscus dryopeorum (Vandel, 1964), Vandel 1964a, Schmalfuss 1979, Schmalfuss 2003, Beron 2016
  Phylum: Arthropoda / Class: Malacostraca / Order: Isopoda / Family: Trichoniscidae
• Henrotiella eubeensis Perreau, 1999, Perreau 1999, Guéorguiev 2007a, Beron 2016
  Phylum: Arthropoda / Class: Insecta / Order: Coleoptera / Family: Leiodidae
• Hystricosphodrus vailatii Casale & Giachino, 2004, Casale & Giachino 2004, Beron 2016, Casale & Giachino 2016
  Phylum: Arthropoda / Class: Insecta / Order: Coleoptera / Family: Carabidae
• Niphargus sp., Beron 2016
  Phylum: Arthropoda / Class: Malacostraca / Order: Amphipoda / Family: Niphargidae
• Pipistrellus kuhlii (Kuhl, 1819), Georgiakakis et al. 2023
  Phylum: Chordata / Class: Mammalia / Order: Chiroptera / Family: Vespertilionidae
• Plusiocampa festai Silvestri, 1932, Condé 1984, Sendra et al. 2020a
  Phylum: Arthropoda / Class: Entognatha / Order: Diplura / Family: Campodeidae
• Rhinolophus blasii Peters, 1866, Georgiakakis et al. 2023
  Phylum: Chordata / Class: Mammalia / Order: Chiroptera / Family: Rhinolophidae
• Rhinolophus euryale Blasius, 1853, Georgiakakis et al. 2023
  Phylum: Chordata / Class: Mammalia / Order: Chiroptera / Family: Rhinolophidae
• Rhinolophus ferrumequinum (Schreber, 1774), Beron 2016, Georgiakakis et al. 2023
  Phylum: Chordata / Class: Mammalia / Order: Chiroptera / Family: Rhinolophidae
• Rhinolophus hipposideros (André, 1797), Beron 2016, Georgiakakis et al. 2023
  Phylum: Chordata / Class: Mammalia / Order: Chiroptera / Family: Rhinolophidae
• Roncus sp., Beron 2016
  Phylum: Arthropoda / Class: Arachnida / Order: Pseudoscorpionida / Family: Neobisiidae
• Tegenaria parietina (Fourcroy, 1785), Naumova et al. 2016
  Phylum: Arthropoda / Class: Arachnida / Order: Araneae / Family: Agelenidae
• Trichoniscus lindbergi Vandel, 1958, Vandel 1964a, Schmalfuss 1979, Schmalfuss et al. 2004, Beron 2016
  Phylum: Arthropoda / Class: Malacostraca / Order: Isopoda / Family: Trichoniscidae

Cave references

• Angelopoulou, E. 1934. The cave of Karystos. To Vouno Greek Mountaineering Club. 1:242–245.
• Anonymous. 1953. Speleological news. To Vouno Greek Mountaineering Club. 1:9-10.
• Skouras, Th. 2000. Caves of Evvoia and Skyros. Bull. Soc. Speleol. Grèce 21:229–261.
• Zervoudakis, I. 1959. Cave of Agia Triada (Karystia). Bull. Soc. Speleol. Grèce 5:76–89.

Species references

• Allegrucci, G., Trucchi, E. & Sbordoni, V. 2011. Tempo and mode of species diversification in Dolichopoda cave crickets (Orthoptera, Rhaphidophoridae). Molecular Phylogenetics and Evolution, 60 (1): 108-121.
• Allegrucci G, Rampini M, Chimenti C, Alexiou S & Di Russo C. 2021. Dolichopoda cave crickets from Peloponnese (Orthoptera, Rhaphidophoridae): molecular and morphological investigations reveal four new species for Greece. The European Zoological Journal, 88(1): 505-524, DOI: 10.1080/24750263.2021.1902005
• Beron, P. 2016. Faune cavernicole de la Grèce. Maison d'édition Est-West., Sofia. pp. 229.
• Blasco-Aróstegui J, Prendini L. 2024. Redescription of Euscorpius studentium Based on Adult Specimens; Updated Classification of Cavernicolous Euscorpiidae; and Review of Cavernicolous Scorpions in the Balkans. Diversity, 2024; 16(12):737. https://doi.org/10.3390/d16120737
• Boudou-Saltet, P. 1971. Les Dolichopodes de Grèce IV. Découverte de la femelle de Dolichopoda insignis. Bull. Soc. Hist. Nat. Toulouse, 107 (3-4): 615-618.
• Boudou-Saltet, P. 1980. Les Dolichopodes (Orth. Rhaph.) de Grèce IX. Une espèce nouvelle en Eubée: Dolichopoda makrykapa. Biol. gallo-hellen., 9 (1): 123-134.
• Boudou-Saltet, P. 1985. Etat actuel de nos connaissances sur les orthoptères cavernicoles de Grèce. Biol. gallo-hellen., 10: 259-263.
• Casale, A. & Giachino, P. M. 2004. Hystricosphodrus vailatii, nuovo genere e nuova specie dell’Isola Eubea (Grecia) (Coleoptera, Carabidae, Sphodrini). Ann. Mus. civ. St. nat. "G. Doria", 96: 341-362.
• Casale A, Giachino PM. . 2016. Nesaeoduvalius (gen. n.) vailatii sp. n., from Euboia island (Eastern Greece) (Coleoptera: Carabidae, Trechinae).. Fragm. Entomol. 48:1–8
• Condé, B. 1984. Diploures Campodeides (Insectes) de Grèce (1ere note). Rev. suisse Zool., 91 (1): 173-201.
• Di Russo, C., Rampini, M. & Cobolli, M. 2014. The cave crickets of Greece: a contribution to the study of Southern Balkan Rhaphidophoridae diversity (Orthoptera), with the description of a new species of Troglophilus Krauss , 1879. Biodiversity Journal, 5 (3): 397-420.
• Fet, V., Soleglad, M. E., Parmakelis, A., Kotsakiozi, P. & Stathi, I. 2014. Two new species of Euscorpius from Euboea Island, Greece (Scorpiones: Euscorpiidae). Arthropoda Selecta, 23 (2): 111-126.
• Fet V., Parmakelis A., Stathi I., Tropea G., Kotsakiozi P., Kardaki L. & Nikolakakis M. 2018. Fauna and zoogeography of scorpions in Greece. In: Sfenthourakis S., Pafilis P., Parmakelis A., Poulakakis N. & Triantis K.A. (Eds) Biogeography and Biodiversity of the Aegean. In honour of Prof. Moysis Mylonas. Broken Hill Publishers Ltd, Nicosia, Cyprus, p. 123-134.
• Georgiakakis P, Kafkaletou Diez A, Salvarina I, Benda P, Billington G, Dietz C, Billington J, Cove D, Davison S, Cooke M, Papadatou E. 2023. The Bats of Greece: An Updated Review of Their Distribution, Ecology and Conservation. Animals. 2023; 13(15):2529. https://doi.org/10.3390/ani13152529
• Guéorguiev, B. 2007. Description of Ceuthophyes enormis nov. sp . from Albania, and notes on the morphology of Henrotiella eubeensis (Coleoptera: Leiodidae: Leptodirini). Boll. Mus. civ. St. nat. Verona, 31: 75-82.
• Kovařik, F. & Beron, P. 2015. A checklist of the scorpions (Arachnida , Scorpiones) in the collections of the National Museum of Natural History (Sofia). Hist. Nat. Bulg., 22: 37-44.
• Naumova, M., Lazarov, S., Petrov, B. & Deltshev, C. 2016. New faunistic data on the cave-dwelling spiders in the Balkan Peninsula (Araneae). Ecologica Montenegrina, 7: 425-438.
• Perreau, M. 1999. Nouveaux genres et nouvelles espèces de Leptodirini (Coleoptera, Leiodidae). Bull. Soc. entomol. Fr., 104 (4): 399-406.
• Rampini, M., Di Russo, C., Taylan, M. S., Gelosa, A. & Cobolli, M. 2012. Four new species of Dolichopoda Bolivar, 1880 from Southern Sporades and Western Turkey (Orthoptera, Rhaphidophoridae, Dolichopodainae). ZooKeys, 201: 43-58.
• Schmalfuss, H. 1979. Revidierte Check-List der Landisopoden (Oniscoidea) Griechenlands. Stuttgart. Beitr. Naturk. Ser. A. (Biol.), 331: 1-42.
• Schmalfuss, H. 2003. World catalog of terrestrial isopods (Isopoda: Oniscidea). Stuttgart. Beitr. Naturk. Ser. A. (Biol.), 654: 1-341.
• Schmalfuss, H., Paragamian, K. & Sfenthourakis, S. 2004. The terrestrial isopods (Isopoda: Oniscidea) of Crete and the surrounding islands. Stuttgart. Beitr. Naturk. Ser. A. (Biol.), 662: 1-74.
• Sendra A., Antić D., Barranco P., Borko Š., Christian E., Delić T., Fadrique F., Faille A., Galli L., Gasparo F., Georgiev D., Giachino P.M., Kováč L., Lukić M., Marcia P., Miculinić K., Nicolosi G., Palero F., Paragamian K., Pérez T., Polak S., Prieto C. 2020. Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1–138. https://doi.org/10.5852/ejt.2020.591
• Vandel, A. 1964. Biospeologie. La Biologie des Animaux Cavernicoles. Gauthier-Villars, Paris. pp. 619.
• Willemse, F. 1984. Catalogue of the Orthoptera of Greece. Fauna Graeciae I. Hellenic Zoological Society. pp. 275.

Citation

Paragamian, K. 2026. Fauna of the cave Spilaio Agias Triadas, Karystos, Karystos, Sterea Ellada. In Paragamian, K., M. Poulinakis, S. Paragamian and I. Nikoloudakis. Cave fauna of Greece database - Hellenic Institute of Speleological Research. Cave data updated 25.10.2023, Accessed 07.03.2026