Spilaio Pelade

Higher Geography
Region: Kentriki Makedonia,
Sub-Region: Serres,
Municipality: Serres,
Closest Settlement: Oinousa
Cave Synonyms
Piladele, Grotte de Saint Jean Ie Prodrome
Cave type
Natural
Protection status
  • GR1260004 - Koryfes Orous Menoikion - Oros Kouskouras - Ypsoma - Spilaio Pelade (Special Area of Conservation): Law 3937/2011 (OGG/Α 60/31.03.2011), Joint Ministerial Decision 50743/2017 (OGG 4432/B/15.12.17). Check site’s Standard Data Form
  • GR1260009 - Koilada Timiou Prodromou-menoikion (Special Protection Area): Law 3937/2011 (OGG/Α 60/31.03.2011), Joint Ministerial Decision 50743/2017 (OGG 4432/B/15.12.17). Check site’s Standard Data Form
Fauna
  • Chthonius orthodactylus (Leach, 1817), Lindberg 1955b, Beron 2016
    Phylum: Arthropoda / Class: Arachnida / Order: Pseudoscorpionida / Family: Chthoniidae
  • Eschatocephalus simplex (Neumann, 1906), Arthur 1956, Beaucournu 1966, Beron 2016
    Phylum: Arthropoda / Class: Arachnida / Order: Ixodida / Family: Ixodidae
  • Eschatocephalus vespertilionis (Koch, 1844), Lindberg 1955d, Arthur 1956
    Phylum: Arthropoda / Class: Arachnida / Order: Ixodida / Family: Ixodidae
  • Eupolybothrus transsylvanicus (Latzel,1882), Stoev 2004, Beron 2016
    Phylum: Arthropoda / Class: Chilopoda / Order: Lithobiomorpha / Family: Lithobiidae
  • Eurocoelotes jurinitschi (Drensky, 1915), Roewer 1959, Brignoli 1979a, Bosmans & Chatzaki 2005, Beron 2016
    Phylum: Arthropoda / Class: Arachnida / Order: Araneae / Family: Agelenidae
  • Laemostenus cimmerius (Fischer von Waldheim, 1823), Coiffait 1955, Lindberg 1955d, Beron 2016
    Phylum: Arthropoda / Class: Insecta / Order: Coleoptera / Family: Carabidae
  • Lithobius crassipes L. Koch, 1862, Beron 2016
    Phylum: Arthropoda / Class: Chilopoda / Order: Lithobiomorpha / Family: Lithobiidae
  • Loxosceles rufescens (Dufour, 1820), Roewer 1959, Brignoli 1979a, Bosmans & Chatzaki 2005, Beron 2016
    Phylum: Arthropoda / Class: Arachnida / Order: Araneae / Family: Sicariidae
  • Maraenobiotus brucei (Richard, 1898), Chappuis 1955, Lindberg 1955d, Beron 2016
    Phylum: Arthropoda / Class: Hexanauplia / Order: Harpacticoida / Family: Canthocamptidae
  • Metellina merianae (Scopoli, 1763), Naumova et al. 2016
    Phylum: Arthropoda / Class: Arachnida / Order: Araneae / Family: Tetragnathidae
  • Miniopterus schreibersii (Kuhl, 1817), Aellen 1955, Lindberg 1955d, Van Laar & Daan 1964, Ondrias 1965, Iliopoulou-Georgudaki 1977, Iliopoulou-Georgudaki & Ondrias 1978, Iliopoulou-Georgudaki 1983b, Hanak et al. 2001, Georgiakakis & Papadatou 2015, Beron 2016, Georgiakakis et al. 2023
    Phylum: Chordata / Class: Mammalia / Order: Chiroptera / Family: Miniopteridae
  • Myotis blythii (Tomes, 1857), Aellen 1955, Lindberg 1955d, Van Laar & Daan 1964, Ondrias 1965, Hanak et al. 2001, Georgiakakis & Papadatou 2015, Beron 2016, Georgiakakis et al. 2023
    Phylum: Chordata / Class: Mammalia / Order: Chiroptera / Family: Vespertilionidae
  • Myotis capaccinii (Bonaparte, 1837), Hanak et al. 2001, Georgiakakis & Papadatou 2015, Georgiakakis et al. 2023
    Phylum: Chordata / Class: Mammalia / Order: Chiroptera / Family: Vespertilionidae
  • Myotis myotis (Borkhausen, 1797), Aellen 1955, Lindberg 1955d, Van Laar & Daan 1964, Ondrias 1965, Iliopoulou-Georgudaki 1977, Iliopoulou-Georgudaki 1983b, Iliopoulou-Georgudaki & Giagia 1984, Hanak et al. 2001, Georgiakakis & Papadatou 2015, Beron 2016, Georgiakakis et al. 2023
    Phylum: Chordata / Class: Mammalia / Order: Chiroptera / Family: Vespertilionidae
  • Nesticus cellulanus (Clerck, 1757), Naumova et al. 2016
    Phylum: Arthropoda / Class: Arachnida / Order: Araneae / Family: Nesticidae
  • Nycteribia latreillei (Leach, 1817), Aellen 1955, Lindberg 1955d, Kock 1974, Beron 2016
    Phylum: Arthropoda / Class: Insecta / Order: Diptera / Family: Nycteribiidae
  • Nycteribia schmidli Schiner, 1853, Aellen 1955, Lindberg 1955d, Kock 1974, Beron 2016
    Phylum: Arthropoda / Class: Insecta / Order: Diptera / Family: Nycteribiidae
  • Nycteribia vexata Westwood, 1835, Aellen 1955, Lindberg 1955d, Kock 1974, Beron 2016
    Phylum: Arthropoda / Class: Insecta / Order: Diptera / Family: Nycteribiidae
  • Paractinolaimus macrolaimus (de Man, 1880), Lindberg 1955b, Beron 2016
    Phylum: Nematoda / Class: Enoplea / Order: Dorylaimida / Family: Actinolaimidae
  • Penicillidia conspicua Speiser, 1901, Aellen 1955, Lindberg 1955d, Kock 1974, Beron 2016
    Phylum: Arthropoda / Class: Insecta / Order: Diptera / Family: Nycteribiidae
  • Penicillidia dufouri (Westwood, 1835), Aellen 1955, Lindberg 1955d, Beron 2016
    Phylum: Arthropoda / Class: Insecta / Order: Diptera / Family: Nycteribiidae
  • Phthiridium biarticulatum Hermann, 1804, Beron 2016
    Phylum: Arthropoda / Class: Insecta / Order: Diptera / Family: Nycteribiidae
  • Plusiocampa lindbergi Condé, 1956, Condé 1956, Beron 2016, Sendra et al. 2020a, Sendra et al. 2025
    Phylum: Arthropoda / Class: Entognatha / Order: Diplura / Family: Campodeidae
  • Porrhomma convexum (Westring, 1851), Naumova et al. 2016
    Phylum: Arthropoda / Class: Arachnida / Order: Araneae / Family: Linyphiidae
  • Rhabditis producta (Schneider, 1866), Lindberg 1955d, Beron 2016
    Phylum: Nematoda / Class: Secernentea / Order: Rhabditida / Family: Rhabditidae
  • Rhinolophus blasii Peters, 1866, Georgiakakis & Papadatou 2015, Georgiakakis et al. 2023
    Phylum: Chordata / Class: Mammalia / Order: Chiroptera / Family: Rhinolophidae
  • Rhinolophus euryale Blasius, 1853, Georgiakakis & Papadatou 2015, Georgiakakis et al. 2023
    Phylum: Chordata / Class: Mammalia / Order: Chiroptera / Family: Rhinolophidae
  • Rhinolophus ferrumequinum (Schreber, 1774), Georgiakakis & Papadatou 2015, Georgiakakis et al. 2023
    Phylum: Chordata / Class: Mammalia / Order: Chiroptera / Family: Rhinolophidae
  • Rhinolophus hipposideros (André, 1797), Georgiakakis & Papadatou 2015, Georgiakakis et al. 2023
    Phylum: Chordata / Class: Mammalia / Order: Chiroptera / Family: Rhinolophidae
  • Rhinolophus mehelyi Matschie, 1901, Lindberg 1955d, Iliopoulou-Georgudaki 1977, Iliopoulou-Georgudaki 1983b, Hanak et al. 2001, Beron 2016, Georgiakakis et al. 2023
    Phylum: Chordata / Class: Mammalia / Order: Chiroptera / Family: Rhinolophidae
  • Sphodrus leucophthalmus (Linnaeus, 1758), Lindberg 1955d, Beron 2016
    Phylum: Arthropoda / Class: Insecta / Order: Coleoptera / Family: Carabidae
  • Spinturnix myoti (Kolenati, 1856), Lindberg 1955d
    Phylum: Arthropoda / Class: Arachnida / Order: Mesostigmata / Family: Spinturnicidae
  • Trechus sp., Beron 2016
    Phylum: Arthropoda / Class: Insecta / Order: Coleoptera / Family: Carabidae
Cave references
  • Lindberg ,K. 1955. Notes sur le grottes de la Grèce. Acta Mus. Macedonici Sci. Nat. 3(2):41-69.
Species references
  • Aellen, V. 1955. Etude d΄ une collection de Nycterobiidae et de Streblidae (Diptera, Pupipara) de la région Palearctique occidentale, particulièrement de la Suisse. Bull. Soc. neuchatel. Sci. nat., 78: 81-104.
  • Arthur, D. R. 1956. The Ixodes ticks of Chiroptera (Ixodoidea: Ixodidae). J. Parasit., 42: 180-196.
  • Beaucournu, J. C. 1966. Sur quelques Ixodoidea (Acarina) palearctiques infeodes aux micro-Chiropteres. Ann. Parasitol. Hum. Comp., 41 (5): 495-502.
  • Beron, P. 2016. Faune cavernicole de la Grèce. Maison d'édition Est-West., Sofia. pp. 229.
  • Bosmans, R. & Chatzaki, M. 2005. A catalogue of the spiders of Greece. A critical review of all spider species cited from Greece with their localities. Newsletter Belg. Arachn. Soc., 20 (2): 1-124.
  • Brignoli, P. M. 1979. Ragni di Grecia XI. Specie nuove o interessanti, cavernicole ed epigee. Rev. suisse Zool., 86 (1): 181-202.
  • Chappuis, P. A. 1955. Notes sur le Copépodes. 18. Un nouveaux Harpacticoides des Pyrénées. 19. Harpacticoides cavernicoles de Grèce, 20. Copépodes Haparcticoides des Iles Pacifique. Notes Biospeol., 10 (1): 89-101.
  • Coiffait, H. 1955. Sur quelques Coléoptères cavernicoles de Grèce. Notes Biospeol., 10 (2): 205-209.
  • Condé, B. 1956. Campodeides cavernicoles de Grèce. Notes biospeol., 11 (1): 7-12.
  • Georgiakakis, P. & Papadatou, E., 2015. Chiroptera, In Papamichael G., Arapis T., Petkidi K., Fytou I & Chatzirvasanis V. 2015. Deliverable 7, C – Phase of Study 7: "Monitoring and evaluation of conservation status mammal species of Community interest in Greece.". Ministry of Environment, Energy and Climatic Change, Athens, scholars Partnership and consultancy firms “ARAPIS THOMAS EYAGGELOY, GEOANALYSI S.A. and PAPACHARISI ALEXANDRA THEODORA ", Athens. pp. 59 (in Greek)
  • Georgiakakis P, Kafkaletou Diez A, Salvarina I, Benda P, Billington G, Dietz C, Billington J, Cove D, Davison S, Cooke M, Papadatou E. 2023. The Bats of Greece: An Updated Review of Their Distribution, Ecology and Conservation. Animals. 2023; 13(15):2529. https://doi.org/10.3390/ani13152529
  • Hanak, V., Benda, P., Ruedi, M., Homcek, I. & Sofianidou, T. 2001. Bats (Mammalia: Chiroptera) of the Eastern Mediterranean. Part 2. New records and review of distribution of bats in Greece. Acta Societatis Zoologicae Bohemicae, 65: 279-346.
  • Iliopoulou-Georgudaki, J. & Giagia, E. B. 1984. Distribution Notes on Myotis myotis (Borkhausen, 1797) (Chiroptera; Vespertilionidae) from Greece, including the Karyotype of Specimens from Lesvos Island. Säugetierk. Mitt., 31: 135-139.
  • Iliopoulou-Georgudaki, J. & Ondrias, J. C. 1978. Population variation in Miniopterus schreibersi from Greece. Biol. gallo-hellen., 7 (1-2): 223-232.
  • Iliopoulou-Georgudaki, J. 1977. Taxonomic study and geographical distribution of Chiroptera in Greece. PhD. Thesis. Faculty of Physics and Mathematics, University of Patra, Patra, 173 pp (in Greek)
  • Iliopoulou-Georgudaki, J. 1983. Bats parasites and their relations to human health. Proceedings of the International Meeting on the show caves and their problems. Publisher, Athens, September 1-3, 1983. pp. 237-243.
  • Kock, D. 1974. Pupipare Dipteren von Säugetieren des nordöstlichen Mittelmeerraumes (Ins.: Diptera). Senckenbergiana biol., 55 (1/3): 87-104.
  • Lindberg, K. 1955. Cyclopides (Crustacés copépodes) de la Grèce II. Fragm. Balc., Skopje, 1 (23): 189-195.
  • Lindberg, K. 1955. Notes sur le grottes de la Grèce. Acta Mus. Maced. Sci. Natur. Skopje 3: 41–69
  • Naumova, M., Lazarov, S., Petrov, B. & Deltshev, C. 2016. New faunistic data on the cave-dwelling spiders in the Balkan Peninsula (Araneae). Ecologica Montenegrina, 7: 425-438.
  • Ondrias, J. C. 1965. Die Säugetiere Griechenlands. Säugetierk. Mitt., 13 (3): 109-127.
  • Roewer, C. F. 1959. Die Araneae, Solifuga und Opiliones der Sammlungen des Herrn Dr. K. Lindberg aus Griechenland, Creta, Anatolien, Iran und Indien. Göteborgs kungl. vet. vitterh. Samh. Handl., 8 (4): 3-47.
  • Sendra A., Antić D., Barranco P., Borko Š., Christian E., Delić T., Fadrique F., Faille A., Galli L., Gasparo F., Georgiev D., Giachino P.M., Kováč L., Lukić M., Marcia P., Miculinić K., Nicolosi G., Palero F., Paragamian K., Pérez T., Polak S., Prieto C. 2020. Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1–138. https://doi.org/10.5852/ejt.2020.591
  • Sendra A, Stoev P, Selfa J 2025. Cave-adapted Plusiocampinae (Diplura, Campodeidae) from the eastern part of the Balkan Peninsula. Deutsche Entomologische Zeitschrift 72(2): 317-340
  • Stoev, P. 2004. Centipedes (Chilopoda) from Greece in the collection of the National Museum of Natural History, Sofia. Hist. Nat. Bulg., 16: 81-88.
  • Van Laar, V. & Daan, S. 1964. On some Chiroptera from Greece. Beaufortia, 120 (10): 158-166.
Citation
Paragamian, K. 2026. Fauna of the cave Spilaio Pelade, Oinousa, Serres, Kentriki Makedonia. In Paragamian, K., M. Poulinakis, S. Paragamian and I. Nikoloudakis. Cave fauna of Greece database - Hellenic Institute of Speleological Research. Cave data updated 08.11.2025, Accessed 08.02.2026