It is not clear whether the Chiropteran records of Niethamer (1974) and Kock (1974) are assigned to the cave of Koufovouno or the cave of Didymoteicho.
Protection status
GR1110005 - Vouna Evrou - spilaia Didymoteichou kai Kefalovounou (Special Area of Conservation): Law 3937/2011 (OGG/Α 60/31.03.2011), Joint Ministerial Decision 50743/2017 (OGG 4432/B/15.12.17). Check site’s Standard Data Form
Benda, P., Ševčík, M., Bego, F., Sachanowicz, K., Spitzenberger, F., Tájek, P., Tájková, P., Uhrin, M. 2019. Bats (Mammalia: Chiroptera) of the Eastern Mediterranean and Middle East. Part 15. The fauna of bats and bat ectoparasites of Albania with a catalogue of bats from the western Balkans in the collection of the National Museum, Prague. Acta Soc. Zool. Bohem. 83: 1–238
Georgiakakis, P. & Papadatou, E., 2015. Chiroptera, In Papamichael G., Arapis T., Petkidi K., Fytou I & Chatzirvasanis V. 2015. Deliverable 7, C – Phase of Study 7: "Monitoring and evaluation of conservation status mammal species of Community interest in Greece.". Ministry of Environment, Energy and Climatic Change, Athens, scholars Partnership and consultancy firms “ARAPIS THOMAS EYAGGELOY, GEOANALYSI S.A. and PAPACHARISI ALEXANDRA THEODORA ", Athens. pp. 59 (in Greek)
Georgiakakis P, Kafkaletou Diez A, Salvarina I, Benda P, Billington G, Dietz C, Billington J, Cove D, Davison S, Cooke M, Papadatou E. 2023. The Bats of Greece: An Updated Review of Their Distribution, Ecology and Conservation. Animals. 2023; 13(15):2529. https://doi.org/10.3390/ani13152529
Hanak, V., Benda, P., Ruedi, M., Homcek, I. & Sofianidou, T. 2001. Bats (Mammalia: Chiroptera) of the Eastern Mediterranean. Part 2. New records and review of distribution of bats in Greece. Acta Societatis Zoologicae Bohemicae, 65: 279-346.
Kock, D. 1974. Pupipare Dipteren von Säugetieren des nordöstlichen Mittelmeerraumes (Ins.: Diptera). Senckenbergiana biol., 55 (1/3): 87-104.
Niethammer, J. 1974. Zur verbreitung und taxonomie griechischer säugetiere. Bonn. Zool. Beitr., 25 (1-3): 28-55.
Citation
Paragamian, K. 2026. Fauna of the cave
Spilaio Didymoteichou, Didymoteicho, Didymoteicho, Anatoliki Makedonia-Thraki. In Paragamian, K., M. Poulinakis, S. Paragamian and I. Nikoloudakis.
Cave fauna of Greece database - Hellenic Institute of Speleological Research.
Cave data updated 15.12.2023,
Accessed 08.02.2026